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Great writeup. Here's a full-text review that contains all the math needed to build a model of this process (2013):

"Quantitative modeling of bacterial chemotaxis: Signal amplification and accurate adaptation, Yuhai Tu"

https://www.ncbi.nlm.nih.gov/pmc/articles/PMC3737589/

The main points are:

* Both receptor cooperativity and accurate adaptation can be described quantitatively by simple mathematical models.

* An integrated model (the “standard model”), which contains both signal amplification and adaptation, is developed to predict responses of it E. coli cells to any time-dependent stimuli quantitatively.

* Exponential ramps induce activity shifts, which depend on the ramp rate through the methylation rate function F(a).

* Responses to oscillatory signals reveal that E. coli computes time-derivative in the low-frequency regime.

* E. coli memorizes the logarithm of the ligand concentration and the Weber-Fetcher law holds in E. coli chemotaxis.

It also goes into cooperative phase transitions in the receptor complexes as a means of signal amplification, using the same model as in Ising ferromagnetic spin-spin interactions in physics.



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